Colour Science & Perception
what the eye settles, and what it cannot
Concept

Afterimage — where it appears

The image left behind after a stimulus is removed, which is an adaptation state relaxing rather than a picture stored anywhere. It has the same time constant as the gain that produced it, which is what joins it to chromatic adaptation.

Named by 6 essays across 2 fields — each of them below, with the objects they name alongside it.

One adapting colour, two answers. The left patch is what was stared at. The middle is the afterimage the cone-gain arithmetic predicts at 15 per cent adaptation; the right is the inverted code values. They are 22.1 ΔE00 apart. The gains that produced the middle patch are 0.95, 1.06, 1.69 on the long, medium and short cone classes — the reciprocal of what each class had been receiving, taken 15 per cent of the way.

An afterimage is an adaptation

The demonstration everybody gives is an inverted image, which is a statement about a file format. Running the receptoral arithmetic instead puts the afterimage of a saturated red sixty degrees of hue away from the inverse — and outside what any display can show.

brain · Appearance An afterimage, as the local pool coming back to equilibrium. The local pool has adapted to the patch and the global pool has not, so the gain change is exactly the local share of a full von Kries change — which is why afterimage's free strength parameter is not free here. The dwell is 20 seconds. The swatches are the predicted appearance of the test surface at four moments. They are predictions of hue and direction; there is no response compression in this model, so the chroma is a ceiling rather than an estimate.

A gain has a time constant

An afterimage and the clock on chromatic adaptation were built in different files from what the last phase said was one mechanism. Joining them removes a free parameter, reproduces both, and predicts a third thing — that two people in one room, at one moment, looking at one patch, do not agree about its colour.

brain · Appearance The eye's own drift, and what it does to every spatial frequency. A pattern of f cycles per degree, drifting across the retina at 0.5 degrees a second, arrives at each receptor at f × 0.5 hertz. The curve is the temporal sensitivity at that rate, against the pattern's spatial frequency. Every frequency the eye can resolve stays above a quarter of the temporal peak, and the band of drift speeds for which that holds is 0.02–0.71 degrees a second — with the measured drift inside it. Faster and the finest detail is carried past 60 hertz, where there is no sensitivity at all.

The eye is never still

A perfectly stabilised retinal image disappears within seconds. What keeps the world there is a drift of about half a degree a second between the microsaccades — fast enough to keep the finest detail modulating and slow enough not to carry it past fusion, in a band whose upper edge is at 0.71 degrees a second.

brain · Appearance Coming back from a bleach, against the clock already measured. A 94 per cent bleach, and the pigment returning at its own time constant of 120 seconds. The lower curve is the site's slow neural adaptation constant, 60 seconds, started from the same place — it is finished while the chemistry is barely half done. Regeneration does not speed up because the light went away: the rate constant is the same one it always was, which is why the recovery is slow while the bleaching was fast.

The slowest clock is chemical

An earlier essay here joined the afterimage to the adaptation clock and named what was still missing — a third gain, upstream of both, in the pigment itself. It is twice as slow as anything measured before it, it leaves a coloured after-tint from a white field, and at steady state it cancels exactly, which is why nobody has ever needed to model it.

eye · Cones Where a pooled gain gives out, against where the eye does. The falling curve is how much of a pattern of each spatial frequency a local adaptation pool of 0.5° can see — and therefore how much of it a settled eye can cancel. It is half gone by 0.37 cycles per degree, which is a feature about 2.7° across. The three marks are the acuity limits of the luminance channel and the two chromatic ones. Every one of them is more than an order of magnitude finer than the pool, which is why a stabilised eye loses the fill of a picture and keeps its outline rather than losing the picture.

What a still eye stops seeing

A stabilised image is said to vanish, and nothing in a temporal filter predicts it — sensitivity at zero frequency is a quarter of the peak, not nothing. Give the adaptation gain a size and the answer falls out — fading is a high-pass filter that switches on over a minute, it takes the fill and leaves the outline, and a patch has to be about two degrees across before it goes at all.

eye · Cones A stabilised grating fades at a rate that depends on which way its bars run. What is left of a 0.3 cycle-per-degree grating, in multiples of its own threshold, as a function of its orientation, at four moments after the image was stabilised on the retina. The first curve is flat to floating point: at the instant the pattern arrives, orientation does not matter, because the filter that carries it has no orientation preference at this frequency. The anisotropy arrives with the fading, reaching ×2.09 after five minutes, and a pool with no axis predicts none of it.

A fading pool has a shape

Giving the local adaptation pool two axes instead of one costs a single parameter and produces a prediction the circular version cannot make — a stabilised grating fades at a rate that depends on which way its bars run. The obvious objection is the oblique effect, and the two act in bands that do not overlap.

eye · Cones

Named alongside it

The objects these essays reach for when they reach for this one.

AdaptationAssertionChromatic adaptationOpponent processingContrast sensitivityEccentricityIndividual variationSpatial frequencyTemporal sensitivityThresholdViewing conditionColour appearance

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