Colour Science & Perception
what the eye settles, and what it cannot
Concept

Visual pigment — where it appears

The light-absorbing molecule in a photoreceptor, whose absorption band decides what that receptor can see. Its peak wavelength varies between people by several nanometres, which is one of the four measured differences a standard observer averages away.

Named by 12 essays across 3 fields — each of them below, with the objects they name alongside it.

The same two surfaces, weighed by each system. A long-wavelength and a short-wavelength surface under D65, with their relative luminance under the photopic curve and under the scotopic one. Under daylight vision the red surface is 1.33 times the blue; under rod vision it is 0.15 times, a reversal by a factor of 9.0. The swatches are the photopic appearance, which is the only one a display can produce: rod vision has no colour, and drawing a guess at it would be an invention.

The eye that has no colour

Rods outnumber cones twenty to one, work alone below a hundredth of a candela, and are absent from the centre of gaze. Between dusk and a lit room both systems run at once, and neither standard curve describes what is happening.

eye · Cones Longitudinal chromatic aberration of the eye, focused at 555 nm. Thibos's chromatic eye, plotted as defocus in dioptres relative to 555 nm. The curve crosses zero exactly once, at the wavelength the eye is accommodating on, and everything else is out of focus — by -1.22 D at 420 nm and 0.50 D at 680 nm. Through a 3 mm pupil that first figure is a blur circle of 12.6 minutes of arc, against a foveal acuity limit of about one. The eye is never in focus across the spectrum and never can be.

Nothing is in focus at both ends

The eye carries about two dioptres of chromatic aberration across the visible band, which is a strong reading prescription. Whatever it is focused on, most of the spectrum is landing somewhere other than the retina — and the cone class that gets the worst of it is the one the retina bothered least to sample.

eye · Cones The L cone's sensitivity at three axial pigment densities. Each curve is normalised to its own peak, so the only difference visible is shape. Raising the density from 0.1 to 0.9 widens the curve from 113.6 to 145.9 nm at half height while the peak stays within 5 nm of where it was. That is Beer–Lambert saturating: at the peak the pigment already absorbs nearly everything, so more of it can only catch more light in the wings.

A cone absorbs its own light

A photopigment's absorbance is a property of a molecule; a cone's sensitivity is that molecule stacked in a column deep enough to absorb most of what arrives. The stacking broadens the curve by thirty-five nanometres, and two observers differing in nothing else disagree about a match that is exact for one of them.

eye · Cones Photons caught by one cone in one integration time, under D65. Each curve is one cone class, counting isomerisations during a 0.1 s integration through a pupil that closes as the light rises. At 1000 cd/m² a long-wavelength cone catches about 11522 and at 0.001 it catches 0.12 — which is where the square root of the count stops being a small correction and starts being the signal. The rate works out at 30 isomerisations per second per troland, inside the published band of 5–50.

Colour goes first in the dark

A cone reports a count, and a count carries the square root of itself as noise. Counting the photons says where colour vision stops — a chromatic difference runs out four hundred times sooner than a lightness difference of the same size — and says just as clearly that in daylight the eye is nowhere near that limit.

eye · Cones Grassmann's four laws, exact — and the two things that break them. For a linear observer every one of the four is exact and the residual is floating point, which is the control that makes the two failures below measurements rather than artefacts. Rods break a cone-metameric match by 23 per cent of a rod excitation at dusk; bleaching breaks it by 0.59 per cent of a cone excitation in the sun. Both are stated as fractions of a receptor's own response, so they can be put on one scale.

The laws that make colour add up

Colorimetry is an integral, and an integral assumes matching is linear. Grassmann's four laws are exact for a linear observer, to floating point — and they fail at both ends of the light range, by two different mechanisms, leaving colorimetry an operating band of three and a bit decades that no standard states.

matching · Gamut One match, and what it says about the observer making it. The anomaloscope: a monochromatic 589 nm yellow set against a mixture of 545 and 670 nm. Only two cone classes respond at those wavelengths, so the match is two equations in two unknowns and has one solution for any observer whose two pigments differ. The bar is the fraction of the accepted band; the mark is the solution. A normal observer accepts 0.7 per cent of the scale; an observer whose two pigments are the same accepts all of it, because their two equations are one equation twice. Nothing here is fitted to clinical data: the pigments are the same template used for every observer here, at stated peaks, and the match is the solution of the linear system.

One match names the observer

A yellow at 589 nanometres set against a mixture of 545 and 670 is two equations in two unknowns. It has one solution for a normal observer, a solution somewhere else for an anomalous one, and no unique solution at all for a dichromat — whose two equations are one equation twice.

matching · Gamut Coming back from a bleach, against the clock already measured. A 94 per cent bleach, and the pigment returning at its own time constant of 120 seconds. The lower curve is the site's slow neural adaptation constant, 60 seconds, started from the same place — it is finished while the chemistry is barely half done. Regeneration does not speed up because the light went away: the rate constant is the same one it always was, which is why the recovery is slow while the bleaching was fast.

The slowest clock is chemical

An earlier essay here joined the afterimage to the adaptation clock and named what was still missing — a third gain, upstream of both, in the pigment itself. It is twice as slow as anything measured before it, it leaves a coloured after-tint from a white field, and at steady state it cancels exactly, which is why nobody has ever needed to model it.

eye · Cones A surface in a room, from the moment the light goes on. How far a blue surface is from the colour it will settle at, second by second, for an observer who walked in from the snow as the lamp was switched on. It starts ΔE00 27 away, is still 6.1 away after a minute — the point at which the eye is conventionally said to have adapted — and does not fall under a unit until 295 seconds.

The room settles after the eye does

Four clocks run in a room and only three of them are in the observer. The slowest is the lamp, which takes four hundred seconds to reach nine tenths of its colour change — so a minute after the light goes on, when the eye is conventionally said to have settled, most of what is left is the lamp, and three quarters of that could not be adapted away by an observer of any speed.

light · Light The share of itself each change leaves behind, and the smallest is inside the eye. The residual as a fraction of the change rather than as a colour difference, which sorts the census differently. At the top is the macular pigment — the filter in front of the central few degrees of one's own retina — leaving 2.4 per cent of itself. It is a fixed transmittance multiplying the light and the white together, which is as close to a pure gain as anything here gets, and it is why nobody notices they have one.

The filters inside the eye

The macular pigment leaves 2.4 per cent of itself after adaptation — the smallest share of anything in this site's census of light changes, and less than half the next smallest. Fifty years of lens yellowing leaves 7.9 per cent, and the difference between the two says what a gain is actually good at.

eye · Cones A template that cannot place a point, fitted three ways. Three rows, one per set of stimuli the pigment template's cone matrix can be fitted over, each listing the three confusion points that matrix implies. The protanope's point wanders from (0.99, 0.20) to (0.76, 0.13) against a measured (0.75, 0.25), and the deuteranope's moves by 22.5 in chromaticity — further than the whole diagram is wide. A copunctal point is where two nearly parallel planes meet, so a template good to a few per cent, which is far more than enough to place a spectrum, is nowhere near enough to place this. It is why the population is built by moving the measured points rather than by deriving them.

A template cannot place a point

This collection's model of an eye is good to a few tenths of a per cent at predicting what a cone catches, which is far more than enough to place a spectrum. Asked where that eye's confusion points are, it puts the protanope's at (0.99, 0.20) against a measured (0.75, 0.25) and the deuteranope's anywhere from (1.1, −0.5) to (−18, 11) depending on which stimuli the fit was made over.

eye · Cones The three cone absorptances at two settings of the pigment peaks. Solid and dashed are the same construction at the two ends of two standard deviations, and the L/M polymorphism on top. The curves are built from one pigment template through its ocular media, which is the same model its population of two hundred eyes is drawn from. The largest difference between the two sets is 8.9 per cent of the peak, and where it sits along the wavelength axis is what decides which stimuli the two observers disagree about — a departure concentrated in the blue is invisible on a sample with no blue in it.

The peaks move the flanks

Shifting a cone's peak wavelength by three nanometres changes its sensitivity at its own maximum by almost nothing and on its flanks by several per cent, because a maximum is flat and a flank is steep. Every display primary sits on a flank, which is why a pigment polymorphism is worth 3.60 ΔE₀₀ under a laser projector.

eye · Cones The same white, matched at six primary widths. At every width the three primaries are solved to match D65 exactly for the reference member; the bands are what the population sees. A broad primary integrates the observer differences over a band and averages them away; a narrow one samples them at a point and passes them straight through. From 40 nm to 2 the ninety-fifth percentile rises from 11.3 to 17.9 ΔE00, monotonically, and the technology has been moving from left to right for thirty years.

A narrow primary buys a disagreement

The observer audit decomposes what a display costs a population. Narrowing the primaries raises the pigment-peak departure monotonically, moving one raises or lowers the macular departure, and the two respond to different design variables — so a wide gamut and an observer-robust display are bought with the same money.

matching · Gamut

Named alongside it

The objects these essays reach for when they reach for this one.

Individual variationCone fundamentalsAdaptationStandard observerLuminanceObserver metamerismMacular pigmentSpectral sensitivityChromatic adaptationColour-matching functionsMetamerismOptical density

All concepts