Colour Science & Perception
what the eye settles, and what it cannot
Concept

Optical density — where it appears

How much pigment lies along a receptor's length, which broadens its sensitivity rather than raising it and differs between people. It is also the reason a matching experiment measures a population rather than a person, since the density varies by a factor of two between eyes.

Named by 6 essays across one field — each of them below, with the objects they name alongside it.

The L cone's sensitivity at three axial pigment densities. Each curve is normalised to its own peak, so the only difference visible is shape. Raising the density from 0.1 to 0.9 widens the curve from 113.6 to 145.9 nm at half height while the peak stays within 5 nm of where it was. That is Beer–Lambert saturating: at the peak the pigment already absorbs nearly everything, so more of it can only catch more light in the wings.

A cone absorbs its own light

A photopigment's absorbance is a property of a molecule; a cone's sensitivity is that molecule stacked in a column deep enough to absorb most of what arrives. The stacking broadens the curve by thirty-five nanometres, and two observers differing in nothing else disagree about a match that is exact for one of them.

eye · Cones Photons caught by one cone in one integration time, under D65. Each curve is one cone class, counting isomerisations during a 0.1 s integration through a pupil that closes as the light rises. At 1000 cd/m² a long-wavelength cone catches about 11522 and at 0.001 it catches 0.12 — which is where the square root of the count stops being a small correction and starts being the signal. The rate works out at 30 isomerisations per second per troland, inside the published band of 5–50.

Colour goes first in the dark

A cone reports a count, and a count carries the square root of itself as noise. Counting the photons says where colour vision stops — a chromatic difference runs out four hundred times sooner than a lightness difference of the same size — and says just as clearly that in daylight the eye is nowhere near that limit.

eye · Cones Coming back from a bleach, against the clock already measured. A 94 per cent bleach, and the pigment returning at its own time constant of 120 seconds. The lower curve is the site's slow neural adaptation constant, 60 seconds, started from the same place — it is finished while the chemistry is barely half done. Regeneration does not speed up because the light went away: the rate constant is the same one it always was, which is why the recovery is slow while the bleaching was fast.

The slowest clock is chemical

An earlier essay here joined the afterimage to the adaptation clock and named what was still missing — a third gain, upstream of both, in the pigment itself. It is twice as slow as anything measured before it, it leaves a coloured after-tint from a white field, and at steady state it cancels exactly, which is why nobody has ever needed to model it.

eye · Cones The three cone absorptances at two settings of the cone optical density. Solid and dashed are the same construction at the two ends of two standard deviations of the reported spread. The curves are built from one pigment template through its ocular media, which is the same model its population of two hundred eyes is drawn from. The largest difference between the two sets is 18.8 per cent of the peak, and where it sits along the wavelength axis is what decides which stimuli the two observers disagree about — a departure concentrated in the blue is invisible on a sample with no blue in it.

A gain is not an observer

Multiply one eye's three cone sensitivities by 1.6, 0.7 and 2.4 and it is not a different eye. The white-point division is that multiplication's inverse, so the two agree exactly — and most of what a cone optical density change does is that multiplication, which is why the largest number in the table of individual variation is the one that matters least.

eye · Cones The three cone absorptances at two settings of the field size. Solid and dashed are the same construction at the two ends of the CIE's own second observer. The curves are built from one pigment template through its ocular media, which is the same model its population of two hundred eyes is drawn from. The largest difference between the two sets is 16.0 per cent of the peak, and where it sits along the wavelength axis is what decides which stimuli the two observers disagree about — a departure concentrated in the blue is invisible on a sample with no blue in it.

A field size is two changes

The CIE publishes two standard observers and the difference between them is usually described as a field size. What actually differs is a macular pigment the light no longer passes through and a cone outer segment the light no longer travels the length of — two changes, in two places, with different signs and different sample dependence.

eye · Cones The three cone absorptances at two settings of the age of the lens. Solid and dashed are the same construction at the two ends of twenty years old against seventy. The curves are built from one pigment template through its ocular media, which is the same model its population of two hundred eyes is drawn from. The largest difference between the two sets is 25.5 per cent of the peak, and where it sits along the wavelength axis is what decides which stimuli the two observers disagree about — a departure concentrated in the blue is invisible on a sample with no blue in it.

The observer has no age

Five of the six arguments in this round are spreads — a population differs about them and the mean is a reasonable summary. The lens is not. Everybody's lens yellows in the same direction at about the same rate, so a standard observer with no age is not an average over a population; it is a snapshot of one moment in every reader's life.

eye · Cones

Named alongside it

The objects these essays reach for when they reach for this one.

Individual variationSelf-screeningCone fundamentalsStandard observerVisual pigmentAdaptationAssertionChromatic adaptationLuminanceMacular pigmentObserver metamerismSpecification

All concepts