Cones — the series
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Three numbers
A spectrum has as many degrees of freedom as anyone cares to give it. The eye reports three. Everything colour science can do, and every way it fails, follows from that one collapse.
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Two spectra, one colour
Metamerism is usually described and almost never demonstrated. It does not have to be — the metameric black space is enormous, so a matching pair can be constructed to order, verified, and then made to come apart by changing the light.
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Why colour is exactly three-dimensional
Matching every wavelength with three primaries requires, for some wavelengths, a negative amount of one of them. That physical awkwardness is why the colour-matching functions were transformed into XYZ, and why the horseshoe is curved.
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The eye that has no colour
Rods outnumber cones twenty to one, work alone below a hundredth of a candela, and are absent from the centre of gaze. Between dusk and a lit room both systems run at once, and neither standard curve describes what is happening.
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Three cones, two axes
The retina does not send three receptor signals down the optic nerve. It sends a sum and two differences, and the reason falls out of the statistics of natural light rather than out of anything the eye intended.
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How fine a colour edge can be
The eye resolves a lightness pattern to about fifty cycles per degree and a red–green one to twelve. Every colour difference here is quoted as though a patch had no size, and the same difference is plainly visible at one scale and gone at another.
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The eye has a shutter
An isoluminant flicker fuses at fifteen hertz and a luminance one at sixty, so a light whose colour changes forty times a second is a steady light of a colour it never emits. And the frequency at which flicker stops being visible is not a property of the eye — it moves twelve and a half hertz for every decade of light.
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Nothing is in focus at both ends
The eye carries about two dioptres of chromatic aberration across the visible band, which is a strong reading prescription. Whatever it is focused on, most of the spectrum is landing somewhere other than the retina — and the cone class that gets the worst of it is the one the retina bothered least to sample.
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A cone absorbs its own light
A photopigment's absorbance is a property of a molecule; a cone's sensitivity is that molecule stacked in a column deep enough to absorb most of what arrives. The stacking broadens the curve by thirty-five nanometres, and two observers differing in nothing else disagree about a match that is exact for one of them.
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A pattern has a direction
Every spatial claim here is a claim about a frequency, and a frequency has no direction in it. Turning a printed screen forty-five degrees makes it exactly twice as quiet with nothing else changed — and the same rotation does nothing at all to a chromatic one.
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Colour stops at the edge of sight
Cone density falls twenty-one-fold between the centre of gaze and ten degrees out, and the three channels give out at three different rates — so a colour difference in the periphery does not merely shrink, it turns. At the exact point of fixation there are no short-wavelength cones at all.
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The mosaic is not the observer
The ratio of long-wavelength to medium-wavelength cones varies between ordinary people by a factor of sixteen. Those people make the same colour matches, to twelve decimal places, and that single fact is the reason a standard observer can exist at all.
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Colour goes first in the dark
A cone reports a count, and a count carries the square root of itself as noise. Counting the photons says where colour vision stops — a chromatic difference runs out four hundred times sooner than a lightness difference of the same size — and says just as clearly that in daylight the eye is nowhere near that limit.
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The drift is a luminance mechanism
The eye's own drift was shown to sit inside a band of speeds that keeps every spatial frequency modulating, and the band was quoted as though it were about vision. Asked about colour, it has no slow edge at all — a stationary chromatic pattern needs no eye movement whatever. And a stabilised chromatic pattern is the first thing to fade.
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The slowest clock is chemical
An earlier essay here joined the afterimage to the adaptation clock and named what was still missing — a third gain, upstream of both, in the pigment itself. It is twice as slow as anything measured before it, it leaves a coloured after-tint from a white field, and at steady state it cancels exactly, which is why nobody has ever needed to model it.
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What a still eye stops seeing
A stabilised image is said to vanish, and nothing in a temporal filter predicts it — sensitivity at zero frequency is a quarter of the peak, not nothing. Give the adaptation gain a size and the answer falls out — fading is a high-pass filter that switches on over a minute, it takes the fill and leaves the outline, and a patch has to be about two degrees across before it goes at all.
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One person is two observers
The macular pigment is a yellow screen over the fovea and nowhere else, so a cone at the centre of gaze and a cone ten degrees away have different colour-matching functions in the same eye. A match made in the middle comes apart at the edge by six units — and fitting one filter to the gap between the CIE's two standard observers gives a density of 0.40 against a measured 0.35.
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The filters inside the eye
The macular pigment leaves 2.4 per cent of itself after adaptation — the smallest share of anything in this site's census of light changes, and less than half the next smallest. Fifty years of lens yellowing leaves 7.9 per cent, and the difference between the two says what a gain is actually good at.
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A fading pool has a shape
Giving the local adaptation pool two axes instead of one costs a single parameter and produces a prediction the circular version cannot make — a stabilised grating fades at a rate that depends on which way its bars run. The obvious objection is the oblique effect, and the two act in bands that do not overlap.
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The eye weights where the light is not
A brightened sheet returns a quarter more light than arrives at 430 nanometres, and it is three tenths of one per cent brighter for it. The luminous efficiency function is 0.017 there against 1.0 in the middle of the band, so the whole effect lands in the blue-yellow axis — brighter than white is a colour claim wearing a brightness word.
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The eye stops at the lens
Neither standard observer is tabulated below 360 nanometres, and the reason is not that the photopigments stop absorbing there. It is that the light never arrives — the cornea and the crystalline lens take it — so the short-wave limit of human colour vision is a piece of optics, it moves by a factor of twenty across a lifetime, and it can be surgically removed.
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The matches do not name the cones
Colour matching is the whole empirical basis of colorimetry, and it fixes the observer's three curves only up to a nonsingular 3×3 — nine numbers that no match, in any quantity, to any precision, can see. One particular choice of those nine is used throughout here, and it was made for a different purpose.
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A confusion point is a missing pigment
The nine numbers colour matching leaves free are fixed by three points on a chromaticity diagram, each of them the place where everything one class of dichromat cannot tell apart converges. Two of the three lie outside the diagram entirely, which is not a defect — a direction in tristimulus space need not correspond to a light.
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A width nobody varied
Five numbers say how much people differ from one another, and every conclusion drawn here about a population rests on them. Each was written down with the range the literature reports beside it, so that a result could be re-read at the pessimistic end. Nothing ever was.
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The three numbers a gain cannot see
Colour matching leaves nine numbers free. Three dichromat confusion points fix six of them and three choices of unit fix the rest — and it turns out that a von Kries gain is exactly blind to those last three. So the dichromat data do not merely constrain an adaptation basis. They determine it, with nothing left over to fit.
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The best axes are not receptors
If the axes that make a von Kries gain work were nearly the cone fundamentals, the published adaptation transforms would be cone-like and their departures would be slack in a fit. They are not. Minimise the residual over all nine free numbers and the winner sits further from the measured dichromat confusion points than any of them.
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Which measurement is worth making
Four things about an eye differ between people, and they have been ranked here by how much of the answers they carry since the population was built. Ranking them by how much doubt they carry gives a different order, and ranking them by which one takes a published claim closest to failing gives a third.
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The rank is the invariance
A von Kries gain cannot see the scale of a row of its basis. That is an identity, proved in a line, and it can be measured instead — as the rank of a second-derivative matrix. Both objectives this collection minimises over the observer's nine free numbers have a Hessian of rank exactly six, and the three directions they cannot see are the three scalings, to a hundredth of a degree.
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A template cannot place a point
This collection's model of an eye is good to a few tenths of a per cent at predicting what a cone catches, which is far more than enough to place a spectrum. Asked where that eye's confusion points are, it puts the protanope's at (0.99, 0.20) against a measured (0.75, 0.25) and the deuteranope's anywhere from (1.1, −0.5) to (−18, 11) depending on which stimuli the fit was made over.
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The surfaces that answer nothing
Five of the hundred and twenty-five test surfaces contribute exactly zero to every number the adaptation census reports — not approximately, exactly — and the reason is the one fact about von Kries adaptation that makes it worth having at all. Counting the set by how much it contributes gives about a hundred members rather than a hundred and twenty-five.
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A point about the pigments that remain
The chromaticity at which a protanope's confusion lines meet does not move at all when the long-wave pigment moves — not slightly, exactly not at all. It moves a great deal when the medium-wave pigment does. A dichromat's confusion point is a fact about the two receptors they have rather than about the one they lack.
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The claim, in nanometres
For four rounds the claim here has been that every published adaptation transform puts the protanope's confusion point outside any real population of eyes, stated in standard deviations of a population whose widths were declared rather than measured. Restated as a pigment displacement it needs no population at all — and the nearest transform asks the medium-wave cone to move thirty per cent of the way to the long-wave one.
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The population rests on a template
Two hundred observers here are built from one formula fitted to microspectrophotometry in 2000. The obvious alternative — the tabulated cone fundamentals — is not available, and the reason is the finding. A tabulated fundamental has no peak wavelength to move, so the moment it is used the population collapses to a single observer.
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Two blurs before the eye's own
A translucent object arrives at the eye already blurred, by a kernel that is a property of the material rather than of the optics. On every material in this collection's table that blur is coarser at ordinary reading distance than the finest detail the eye can resolve — so the softness of marble or skin or wax is not a failure of vision, it is the object.
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A template is mostly its tail
Four pigment templates matched to the same width at the same peak, differing only in which side of the peak their half-maximum reaches further. Ordered by that one number, the observers they build are ordered by how far they sit from the standard one — and the two with the tail on the wrong side cost two and four times what either real nomogram costs.
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The band below four hundred
The pigment template every observer here is built from carries a second, smaller absorption band in the ultraviolet, published at 0.26 of the main one. Dialling it from nothing to twice that moves the median observer monotonically away from the standard one, with no interior optimum — which is what a physical constant looks like when nothing downstream is pulling it anywhere.
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The third factor is a construction
Every figure in this collection names its observer, which was the whole point. None of them says what an observer is made of. Three curves are not a measurement of the eye; they are a projection of one, with a field size, an age, a macular density, three peak wavelengths and a luminance constraint inside them.
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A neutral is everyone's colour
Two eyes differing by fifty years of lens yellowing, by a factor of three in macular pigment and by six nanometres of long-wavelength peak agree about a grey card to four parts in ten thousand billion. The agreement is an identity rather than a coincidence, and it says exactly what an observer disagreement is a disagreement about.
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A gain is not an observer
Multiply one eye's three cone sensitivities by 1.6, 0.7 and 2.4 and it is not a different eye. The white-point division is that multiplication's inverse, so the two agree exactly — and most of what a cone optical density change does is that multiplication, which is why the largest number in the table of individual variation is the one that matters least.
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Three curves for one space
Rotate a set of colour-matching functions by an arbitrary invertible matrix, undo the rotation at the end, and the computed colour is identical to eight parts in a thousand million million. An observer is a three-dimensional subspace, not a set of curves, and the literature keeps reopening a question that is a theorem.
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The lens is worst under tungsten
An ageing lens costs 4.38 ΔE₀₀ under a tungsten lamp and 2.20 under a fluorescent tube. The pigment peaks cost 2.84 under a three-emitter LED and 1.87 under the same tube. Six departures across six lights do not form a single ordering, which is what a pairing looks like and what a dominant term would not.
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The identity is in the eye's own coordinates
Two conditions in this round are exact — a gain on each cone is not a different observer, and three curves for one space are one observer. Imposed in a published cone space rather than the eye's own they leave 13.0 and 8.11 ΔE₀₀ standing. The identities belong to the physiology and every arithmetic in use works somewhere else.
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The ranking is not stable
On a red pigment under daylight the six observer departures run from 2.38 down to 1.20 ΔE₀₀. Over forty-two surfaces two of them change places, the top two separate, and every one spans between a factor of ten and a factor of thirty-five. A chart of six bars is a chart of one example.
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A field size is two changes
The CIE publishes two standard observers and the difference between them is usually described as a field size. What actually differs is a macular pigment the light no longer passes through and a cone outer segment the light no longer travels the length of — two changes, in two places, with different signs and different sample dependence.
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The observer has no age
Five of the six arguments in this round are spreads — a population differs about them and the mean is a reasonable summary. The lens is not. Everybody's lens yellows in the same direction at about the same rate, so a standard observer with no age is not an average over a population; it is a snapshot of one moment in every reader's life.
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The rods are a fourth curve
Colorimetry has three numbers and a rod signal is a fourth. It cannot be absorbed by a gain, it cannot be removed by a white point, and adding a tenth of one to a three-curve observer costs 1.60 ΔE₀₀ — the narrowest distribution of the six departures, because an addition behaves quite differently from a filter.
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The macular is a band, not a filter
The lens absorbs everything below 500 nanometres with a long tail; the macular pigment absorbs forty nanometres either side of 460 and nothing else. The two have similar sizes and completely different distributions, and the reason is that one is broad and one is narrow — which is what decides whether a sample is affected at all.
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The peaks move the flanks
Shifting a cone's peak wavelength by three nanometres changes its sensitivity at its own maximum by almost nothing and on its flanks by several per cent, because a maximum is flat and a flank is steep. Every display primary sits on a flank, which is why a pigment polymorphism is worth 3.60 ΔE₀₀ under a laser projector.
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Two observers and one metamer
An observer departure is invisible on a single sample compared with nothing. It becomes a disagreement the moment two spectra are being asked to match, because a match is an identity between three integrals and a different observer takes different integrals. Everything in this round is a statement about pairs wearing a single sample's clothes.
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The rods' route is priced by the lamp
A rod signal in a dim room disturbs a colour match, and how much depends on which of the cone pathways it reaches — a weight the physiology leaves uncertain, especially for the blue–yellow pathway. Under daylight the uncertainty is nearly free: a rod signal that skips the S pathway costs 0.90 at the median surface against 1.03 for one that enters all three. Under a phosphor white LED it is worth a factor of 2.75, 0.45 against 1.24. What decides it is one number per lamp: how large the rod signal is compared with each cone class's own catch of the light.
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The reference lamp must not move
To measure an uncertain weight, use the condition in which the answer depends on it most. That is right about half of an asymmetric colour match and exactly wrong about the other half: a match measures a difference of two displacements, and a reference field that also moves with the weight cancels the signal the test field carries. Daylight is the least sensitive of five lamps and belongs in every one of the three best pairs — 75 settings against a phosphor LED, 2,804 against the pair of lamps the principle as stated would have chosen.
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A rod signal has no natural size
An older lens absorbs where the S cones are sensitive, so it should make the uncertain rod-to-S-cone weight cheaper. Measured, the rod signal's catch of a phosphor LED as a share of the S cones' own catch more than doubles from twenty to seventy-five — and the share the model actually uses falls by a fifth. The two differ by the S cone's peak absorptance, which the lens takes 60 per cent of, and which entered the model as a normalisation rather than as a claim.
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One field keeps what two rooms divide out
An asymmetric match can measure how strongly the rods feed the blue-yellow pathway, but set with the observer adapted to each lamp in turn it needs seventy-five settings on the best pair of lamps and hours of waiting between them. Putting the two lamps on the two halves of one field was proposed as the quick version, at the cost of a weaker signal. The signal is not weaker. Under one shared adaptation it is three times stronger for daylight against a white LED, and the best pair needs six settings. The adaptation that makes the slow version slow is also what was dividing the rods' contribution out of each half.