Concept

Photopigment — where it appears

The light-absorbing molecule inside a receptor, an opsin bound to a chromophore, whose absorption spectrum sets the shape of that receptor's sensitivity. Light bleaches it and the eye remakes it chemically over minutes, so how much is present depends on the light of the last few minutes.

Named by 3 essays across one field — each of them below, with the objects they name alongside it.

What reaches the retina, and why the observer's table stops at 360 nanometres. The transmittance of the eye's own optics across the short-wave band, at three ages, with the brightener's absorption shaded underneath. The upper curve is an eye whose lens has been removed — the cornea alone, opaque below about 295 nanometres and transparent above it. The photopigments absorb perfectly well in this band; what stops the light is a piece of optics in front of them, which is why the short-wave limit of colour vision moves with age and can be removed surgically. A twenty-year-old receives 21 times as much of the band a brightener works in as a seventy-year-old does.

The eye stops at the lens

Neither standard observer is tabulated below 360 nanometres, and the reason is not that the photopigments stop absorbing there. It is that the light never arrives — the cornea and the crystalline lens take it — so the short-wave limit of human colour vision is a piece of optics, it moves by a factor of twenty across a lifetime, and it can be surgically removed.

eye · Cones
What the rods cost a match, by where their signal enters and under which lamp. For five lights, the median colour difference over forty-two surfaces between the reference observer and the same observer with a rod signal a tenth of each cone's peak added — into all three cone channels, into the long- and middle-wavelength channels only, or into the short-wavelength channel only. Under daylight the first two are 1.03 and 0.90: whether the rods reach the S pathway hardly matters. Under a phosphor white LED they are 1.24 and 0.45, a factor of 2.75, and the S-only route alone costs 0.96.

The rods' route is priced by the lamp

A rod signal in a dim room disturbs a colour match, and how much depends on which of the cone pathways it reaches — a weight the physiology leaves uncertain, especially for the blue–yellow pathway. Under daylight the uncertainty is nearly free: a rod signal that skips the S pathway costs 0.90 at the median surface against 1.03 for one that enters all three. Under a phosphor white LED it is worth a factor of 2.75, 0.45 against 1.24. What decides it is one number per lamp: how large the rod signal is compared with each cone class's own catch of the light.

eye · Cones
Two ways to ask how much rod signal an older eye has. The rod signal's catch of each lamp divided by the S cones' own catch, against the observer's age, for five lamps — drawn twice. The upper curves take the rod signal at a fixed absolute size, and every one of them roughly doubles from twenty to seventy-five: an older lens cuts the blue before the S cones see it and the rods, peaking further into the green, lose much less. The lower curves take the rod signal at a tenth of each cone's own peak absorptance, which is the model's own definition, and they barely move at all. Nothing about the retina differs between the two; only the normalisation does.

A rod signal has no natural size

An older lens absorbs where the S cones are sensitive, so it should make the uncertain rod-to-S-cone weight cheaper. Measured, the rod signal's catch of a phosphor LED as a share of the S cones' own catch more than doubles from twenty to seventy-five — and the share the model actually uses falls by a fifth. The two differ by the S cone's peak absorptance, which the lens takes 60 per cent of, and which entered the model as a normalisation rather than as a claim.

eye · Cones

Named alongside it

The objects these essays reach for when they reach for this one.

Cone fundamentalsLens yellowingMesopicModelling assumptionObserver variabilityRodsWhite LEDIndividual differencesIndividual variationIntegrationMacular pigmentMeasurement condition

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